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Form 13F-HR Portolan Capital Managem For: Mar 31

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May 16, 2022 2:47 PM EDT

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F COVER PAGE
OMB APPROVAL
OMB Number: 3235-0006
Expires: July 31, 2015
Estimated average burden
hours per response: 23.8
Report for the Calendar Year or Quarter Ended: 03-31-2022
Check here if Amendment   Amendment Number:
This Amendment (Check only one.):   is a restatement.
  adds new holdings entries.
Institutional Investment Manager Filing this Report:
Name: Portolan Capital Management, LLC
Address: 2 International Place, 26th Floor
Boston , MA02110
Form 13F File Number: 028-11602

The institutional investment manager filing this report and the person by whom it is signed hereby represent that the person signing the report is authorized to submit it, that all information contained herein is true, correct and complete, and that it is understood that all required items, statements, schedules, lists, and tables, are considered integral parts of this form.

Person Signing this Report on Behalf of Reporting Manager:
Name: Betsy Flaherty
Title: Chief Compliance Officer
Phone: 617-753-6400
Signature, Place, and Date of Signing:
/s/ Betsy Flaherty Boston , MA 05-16-2022
[Signature] [City, State] [Date]
Report Type (Check only one.):
X 13F HOLDINGS REPORT. (Check here if all holdings of this reporting manager are reported in this report.)
  13F NOTICE. (Check here if no holdings reported are in this report, and all holdings are reported by other reporting manager(s).)
  13F COMBINATION REPORT. (Check here if a portion of the holdings for this reporting manager are reported in this report and a portion are reported by other reporting manager(s).)
Form 13F Summary Page
Report Summary:
Number of Other Included Managers: 0
Form 13F Information Table Entry Total: 136
Form 13F Information Table Value Total: 1,254,638
(thousands)
List of Other Managers Reporting for this Manager:
[If there are no entries in this list, omit this section.]
Form 13F File Number Name

List of Other Included Managers:
Provide a numbered list of the name(s) and Form 13F file number(s) of all institutional investment managers with respect to which this report is filed, other than the manager filing this report.
[If there are no entries in this list, state “NONE” and omit the column headings and list entries.]
No. Form 13F File Number Name

The Securities and Exchange Commission has not necessarily reviewed the information in this filing and has not determined if it is accurate and complete.
The reader should not assume that the information is accurate and complete.

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
FORM 13F

FORM 13F INFORMATION TABLE
OMB APPROVAL
OMB Number: 3235-0006
Expires: July 31, 2015
Estimated average burden
hours per response: 23.8
COLUMN 1 COLUMN 2 COLUMN 3 COLUMN 4 COLUMN 5 COLUMN 6 COLUMN 7 COLUMN 8
VALUE SHRS OR SH/ PUT/ INVESTMENT OTHER VOTING AUTHORITY
NAME OF ISSUER TITLE OF CLASS CUSIP (x$1000) PRN AMT PRN CALL DISCRETION MANAGER SOLE SHARED NONE
ABIOMED, INC. COM 003654100 2,370 7,154 SH SOLE 7,154 0 0
ACADEMY SPORTS & OUTDOORS, INC. COM 00402L107 840 21,332 SH SOLE 21,332 0 0
AGCO CORPORATION COM 001084102 9,220 63,135 SH SOLE 63,135 0 0
ALBEMARLE CORPORATION COM 012653101 5,791 26,187 SH SOLE 26,187 0 0
ALPHABET, INC. COM 02079K107 13,256 4,746 SH SOLE 4,746 0 0
ALTA EQUIPMENT GROUP, INC. COM 02128L106 19,465 1,574,848 SH SOLE 1,574,848 0 0
AMAZON.COM, INC. COM 023135106 10,839 3,325 SH SOLE 3,325 0 0
AMEDISYS, INC. COM 023436108 2,861 16,607 SH SOLE 16,607 0 0
ANDERSONS, INC. COM 034164103 15,102 300,482 SH SOLE 300,482 0 0
APPLIED GENETIC TECHNOLOGIES CORPORATION COM 03820J100 1,718 1,605,599 SH SOLE 1,605,599 0 0
ARCBEST CORPORATION COM 03937C105 3,743 46,501 SH SOLE 46,501 0 0
ARCHER-DANIELS-MIDLAND COMPANY COM 039483102 14,035 155,496 SH SOLE 155,496 0 0
ATLAS AIR WORLDWIDE HOLDINGS COM 049164205 2,558 29,613 SH SOLE 29,613 0 0
AVID BIOSERVICES, INC. COM 05368M106 4,464 219,125 SH SOLE 219,125 0 0
AVID TECHNOLOGY, INC. COM 05367P100 25,781 739,345 SH SOLE 739,345 0 0
BABCOCK & WILCOX ENTERPRISES, INC. COM 05614L209 9,268 1,135,782 SH SOLE 1,135,782 0 0
BANDWIDTH, INC. COM 05988J103 33,021 1,019,493 SH SOLE 1,019,493 0 0
BIO-RAD LABORATORIES, INC. COM 090572207 19,320 34,303 SH SOLE 34,303 0 0
BLUELINX HOLDINGS, INC. COM 09624H208 19,387 269,711 SH SOLE 269,711 0 0
BTRS HOLDINGS, INC. COM 11778X104 1,036 138,559 SH SOLE 138,559 0 0
BUCKLE, INC. COM 118440106 3,168 95,897 SH SOLE 95,897 0 0
BUNGE LIMITED COM G16962105 10,341 93,325 SH SOLE 93,325 0 0
CAPRI HOLDINGS LIMITED COM G1890L107 11,720 228,062 SH SOLE 228,062 0 0
CASELLA WASTE SYSTEMS, INC. COM 147448104 1,499 17,097 SH SOLE 17,097 0 0
CATALENT, INC. COM 148806102 9,644 86,961 SH SOLE 86,961 0 0
CF INDUSTRIES HOLDINGS, INC. COM 125269100 3,735 36,239 SH SOLE 36,239 0 0
CHAMPIONX CORPORATION COM 15872M104 5,717 233,550 SH SOLE 233,550 0 0
CHARAH SOLUTIONS, INC. COM 15957P105 17,924 3,591,972 SH SOLE 3,591,972 0 0
CHESAPEAKE ENERGY CORPORATION COM 165167735 2,965 34,077 SH SOLE 34,077 0 0
CHILDREN'S PLACE, INC. (THE) COM 168905107 3,319 67,330 SH SOLE 67,330 0 0
COMMERCIAL VEHICLE GROUP, INC. COM 202608105 323 38,212 SH SOLE 38,212 0 0
CROCS, INC. COM 227046109 26,110 341,755 SH SOLE 341,755 0 0
CUSTOM TRUCK ONE SOURCE, INC. COM 23204X103 12,153 1,448,509 SH SOLE 1,448,509 0 0
CUSTOMERS BANCORP, INC. COM 23204G100 25,492 488,908 SH SOLE 488,908 0 0
CYBEROPTICS CORPORATION COM 232517102 6,243 153,844 SH SOLE 153,844 0 0
DANAOS CORPORATION COM Y1968P121 1,426 13,905 SH SOLE 13,905 0 0
DASEKE, INC. COM 23753F107 26,511 2,632,670 SH SOLE 2,632,670 0 0
DESTINATION XL GROUP, INC. COM 25065K104 1,307 267,843 SH SOLE 267,843 0 0
DHT HOLDINGS, INC. COM Y2065G121 2,994 516,213 SH SOLE 516,213 0 0
DIGITAL TURBINE, INC. COM 25400W102 28,467 649,789 SH SOLE 649,789 0 0
DOMO, INC. COM 257554105 37,610 743,725 SH SOLE 743,725 0 0
E.L.F. BEAUTY, INC. COM 26856L103 14,372 556,404 SH SOLE 556,404 0 0
ELANCO ANIMAL HEALTH INCORPORATED COM 28414H103 2,682 102,813 SH SOLE 102,813 0 0
ENCORE CAPITAL GROUP, INC. COM 292554102 1,877 29,923 SH SOLE 29,923 0 0
ENOVA INTERNATIONAL, INC. COM 29357K103 4,580 120,614 SH SOLE 120,614 0 0
EURONET WORLDWIDE, INC. COM 298736109 6,176 47,453 SH SOLE 47,453 0 0
F45 TRAINING HOLDINGS, INC. COM 30322L101 380 35,547 SH SOLE 35,547 0 0
F5, INC. COM 315616102 10,225 48,937 SH SOLE 48,937 0 0
FERROGLOBE PLC COM G33856108 5,182 673,015 SH SOLE 673,015 0 0
FIDELITY NATIONAL INFORMATION SERVICES, INC. COM 31620M106 6,287 62,606 SH SOLE 62,606 0 0
FULL HOUSE RESORTS COM 359678109 11,286 1,174,411 SH SOLE 1,174,411 0 0
GENESCO, INC. COM 371532102 10,372 163,059 SH SOLE 163,059 0 0
GFL ENVIRONMENTAL, INC. COM 36168Q104 10,649 327,257 SH SOLE 327,257 0 0
GOLAR LNG LIMITED COM G9456A100 8,305 335,159 SH SOLE 335,159 0 0
GREEN PLAINS, INC. COM 393222104 11,551 372,502 SH SOLE 372,502 0 0
GTY TECHNOLOGY HOLDINGS, INC. COM 362409104 4,512 1,397,039 SH SOLE 1,397,039 0 0
GULFPORT ENERGY CORPORATION COM 402635502 1,332 14,833 SH SOLE 14,833 0 0
H&E EQUIPMENT SERVICES, INC. COM 404030108 7,753 178,139 SH SOLE 178,139 0 0
HARVARD BIOSCIENCE, INC. COM 416906105 15,691 2,526,789 SH SOLE 2,526,789 0 0
HCI GROUP, INC. COM 40416E103 5,815 85,283 SH SOLE 85,283 0 0
HEALTH CATALYST, INC. COM 42225T107 7,249 277,407 SH SOLE 277,407 0 0
HEALTHEQUITY, INC. COM 42226A107 8,650 128,261 SH SOLE 128,261 0 0
HIMAX TECHNOLOGIES, INC. COM 43289P106 3,014 277,528 SH SOLE 277,528 0 0
HUDSON PACIFIC PROPERTIES, INC. COM 444097109 6,763 243,702 SH SOLE 243,702 0 0
I3 VERTICALS, INC. COM 46571Y107 7,573 271,833 SH SOLE 271,833 0 0
IAC/INTERACTIVECORP. COM 44891N208 723 7,206 SH SOLE 7,206 0 0
ICAD, INC. COM 44934S206 4,041 905,984 SH SOLE 905,984 0 0
IDENTIV, INC. COM 45170X205 32,820 2,029,694 SH SOLE 2,029,694 0 0
IMMERSION CORPORATION COM 452521107 7,029 1,264,245 SH SOLE 1,264,245 0 0
INOTIV, INC. COM 45783Q100 10,236 390,988 SH SOLE 390,988 0 0
INSPIRED ENTERTAINMENT, INC. COM 45782N108 6,024 489,720 SH SOLE 489,720 0 0
KALEYRA, INC. COM 483379103 9,915 1,658,066 SH SOLE 1,658,066 0 0
KURA SUSHI USA, INC. COM 501270102 18,482 335,127 SH SOLE 335,127 0 0
LANTRONIX, INC. COM 516548203 334 50,000 SH SOLE 50,000 0 0
LAREDO PETROLEUM, INC. COM 516806205 16,869 213,150 SH SOLE 213,150 0 0
LEIDOS HLDGS, INC. COM 525327102 10,342 95,745 SH SOLE 95,745 0 0
LESLIE'S, INC. COM 527064109 3,829 197,787 SH SOLE 197,787 0 0
LIVEPERSON, INC. COM 538146101 13,705 561,227 SH SOLE 561,227 0 0
MAGNACHIP SEMICONDUCTOR CORPORATION COM 55933J203 2,875 170,020 SH SOLE 170,020 0 0
MATCH GROUP, INC. COM 57667L107 805 7,406 SH SOLE 7,406 0 0
MATERION CORPORATION COM 576690101 3,337 38,922 SH SOLE 38,922 0 0
MAXLINEAR, INC. COM 57776J100 2,808 48,121 SH SOLE 48,121 0 0
META PLATFORMS, INC. COM 30303M102 8,715 39,191 SH SOLE 39,191 0 0
MGIC INVESTMENT CORPORATION COM 552848103 3,279 241,990 SH SOLE 241,990 0 0
MGP INGREDIENTS, INC. COM 55303J106 7,369 86,101 SH SOLE 86,101 0 0
MICROSOFT CORPORATION COM 594918104 4,210 13,656 SH SOLE 13,656 0 0
NMI HOLDINGS, INC. COM 629209305 4,400 213,405 SH SOLE 213,405 0 0
NOODLES & COMPANY COM 65540B105 1,659 277,845 SH SOLE 277,845 0 0
NORTHERN OIL AND GAS, INC. COM 665531307 13,469 477,799 SH SOLE 477,799 0 0
NUTRIEN LTD. COM 67077M108 3,620 34,815 SH SOLE 34,815 0 0
OLIN CORPORATION COM 680665205 7,872 150,583 SH SOLE 150,583 0 0
PARTY CITY HOLDCO, INC. COM 702149105 1,790 500,000 SH SOLE 500,000 0 0
PDC ENERGY, INC. COM 69327R101 3,469 47,723 SH SOLE 47,723 0 0
PETIQ, INC. COM 71639T106 9,476 388,358 SH SOLE 388,358 0 0
PFSWEB, INC. COM 717098206 313 27,363 SH SOLE 27,363 0 0
PORCH GROUP, INC. COM 733245104 5,344 769,499 SH SOLE 769,499 0 0
QUALYS, INC. COM 74758T303 5,580 39,186 SH SOLE 39,186 0 0
QUINSTREET, INC. COM 74874Q100 9,994 861,538 SH SOLE 861,538 0 0
R1 RCM, INC. COM 749397105 8,828 329,901 SH SOLE 329,901 0 0
RESOLUTE FOREST PRODUCTS, INC. COM 76117W109 6,852 530,772 SH SOLE 530,772 0 0
REX AMERICAN RESOURCES CORPORATION COM 761624105 10,017 100,570 SH SOLE 100,570 0 0
RH COM 74967X103 652 1,998 SH SOLE 1,998 0 0
SANDRIDGE ENERGY, INC. COM 80007P869 5,787 361,211 SH SOLE 361,211 0 0
SCIENCE APPLICATIONS INTERNATIONAL CORPORATION COM 808625107 4,734 51,363 SH SOLE 51,363 0 0
SHYFT GROUP, INC. (THE) COM 825698103 918 25,416 SH SOLE 25,416 0 0
SIBANYE STILLWATER LIMITED COM 82575P107 7,413 456,761 SH SOLE 456,761 0 0
SIGNIFY HEALTH, INC. COM 82671G100 6,129 337,693 SH SOLE 337,693 0 0
SKECHERS USA, INC. COM 830566105 10,567 259,241 SH SOLE 259,241 0 0
SKYLINE CHAMPION CORPORATION COM 830830105 3,026 55,139 SH SOLE 55,139 0 0
SONY GROUP CORPORATION COM 835699307 13,267 129,172 SH SOLE 129,172 0 0
STAAR SURGICAL COMPANY COM 852312305 8,934 111,798 SH SOLE 111,798 0 0
STEWART INFORMATION SERVICES CORPORATION COM 860372101 4,314 71,184 SH SOLE 71,184 0 0
STRATASYS LTD COM M85548101 26,940 1,061,067 SH SOLE 1,061,067 0 0
SUNOPTA, INC. COM 8676EP108 7,349 1,463,962 SH SOLE 1,463,962 0 0
SVB FINANCIAL GROUP COM 78486Q101 4,036 7,214 SH SOLE 7,214 0 0
SYNOVUS FINANCIAL CORP. COM 87161C501 7,195 146,838 SH SOLE 146,838 0 0
TAKE TWO INTERACTIVE SOFTWARE, INC. COM 874054109 21,110 137,311 SH SOLE 137,311 0 0
TALOS ENERGY, INC. COM 87484T108 9,172 580,868 SH SOLE 580,868 0 0
TAYLOR MORRISON HOME CORP. COM 87724P106 2,971 109,154 SH SOLE 109,154 0 0
TECHNIPFMC PLC COM G87110105 5,165 666,424 SH SOLE 666,424 0 0
TECK RESOURCES LIMITED COM 878742204 10,244 253,627 SH SOLE 253,627 0 0
TECNOGLASS, INC. COM G87264100 22,852 905,373 SH SOLE 905,373 0 0
TENABLE HOLDINGS, INC. COM 88025T102 11,321 195,897 SH SOLE 195,897 0 0
TEXTAINER GROUP HOLDINGS COM G8766E109 19,851 521,432 SH SOLE 521,432 0 0
THERMO FISHER SCIENTIFIC, INC. COM 883556102 12,282 20,794 SH SOLE 20,794 0 0
TRANSMEDICS GROUP, INC. COM 89377M109 10,233 379,859 SH SOLE 379,859 0 0
TRAVELCENTERS OF AMERICA, INC. COM 89421B109 25,113 584,556 SH SOLE 584,556 0 0
TRIPADVISOR, INC. COM 896945201 4,094 150,955 SH SOLE 150,955 0 0
TRITON INTERNATIONAL LIMITED COM G9078F107 15,199 216,566 SH SOLE 216,566 0 0
TWILIO, INC. COM 90138F102 8,359 50,721 SH SOLE 50,721 0 0
TWITTER, INC. COM 90184L102 14,440 373,234 SH SOLE 373,234 0 0
VALERO ENERGY CORPORATION COM 91913Y100 2,962 29,168 SH SOLE 29,168 0 0
VISTRA CORP. COM 92840M102 29,054 1,249,640 SH SOLE 1,249,640 0 0
WEST FRASER TIMBER CO. LTD. COM 952845105 13,537 164,225 SH SOLE 164,225 0 0
WORKDAY, INC. COM 98138H101 4,047 16,899 SH SOLE 16,899 0 0
ZIM INTEGRATED SHIPPING SERVICES LTD. COM M9T951109 15,917 218,911 SH SOLE 218,911 0 0

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